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That doesn't really make a whole lot of sense. How can he lead the team in tackles but "struggled against the run"?
They were ranked 32 against the run. A player can get a lot of tackles and be bad against the run. Tackles includes receivers and TEs and pass-catching RBs. The team was thin at LB, so some mix of LBs will get the tackles. It ended up being him. Usually LBs who are bad against the run don't rack up tackles, because the coaches have other options at the position. But all they had were developing LBs.